1 Information and Computational Sciences, University of Dundee, Errol Rd, Invergowrie, Dundee, DD2 5DA, UK; Dundee Effector Consortium, University of Dundee, Errol Rd, Invergowrie, Dundee, DD2 5DA, UK; Centre for Human and Animal Pathogens in the Environment, University of Dundee, Errol Rd, Invergowrie, Dundee, DD2 5DA, UK.